Hi Prof. I know I'm wasting my time but I couldn't resist it.
You should know me well enough to realise that I enjoy playing Devil's Advocate. I don't know the truth & doubt it will ever be uncovered now. Mind you, the article I mentioned is extremely interesting & proves that the information on the "Meredith Effect" was available & documented in 1935 if anyone was prepared to read it. From what I can make out, 2 prototypes of the XP-46 were ordered on September 29, 1939 & the first flight did not take place until September 29, 1941, exactly 2 years later. The Mustang was already in production & being evaluated by the RAF by then. The Napier Heston in Charlie's photo is another example of the ventral positioned radiator scoop which was certainly not unique, not in Europe anyway.
However, from what "Lee" Atwood explains in the article the Heston would not have benefitted from the Meredith Effect. This is caused by the design & shape of the radiator scoop itself. The important thing seems to be that the front part of the scoop is clear of the fuselage as it is on the P-51D & not directly bolted on or faired into the fuselage itself as on the Heston & so many other examples. This is clearly shown on this photo I took of the immaculate P-51D "Jumpin' Jacques" during a fly-in at Abingdon earlier this year.

I don't think the all-important gap between the air intake & the fuselage was so marked on the original P-51 & this was developed over a period of time. It would have been mainly a matter of trial & error as the results couldn't be verified in the wind-tunnels available at the time. The full extent of the air scoop's effect on perfomance was not fully appreciated until after WWII was over.
I think any similarities between the P-40 & P-51 might be exaggerated by the fact that US-designed fighters were invariably radial engined so wouldn't require a radiator at all. (The Curtiss P-75 Hawk, direct predecessor of the P-40, was powered by the Pratt & Whitney Twin Wasp.) These are the only ones I can think of that had inline water-cooled engines, both originally having the same engine. The photo in your previous reply only serves to remind me of the basic differences between the two types.

PS. It didn't strike me until recently how similar the air intake on the F-16 is to the Mustang air scoop. This might explain its remarkable performance.